I think it was a few things. There was the 'Ogay' sprite, as a joke against Olay. There was a 'dev' sprite of text with a bad joke that was never cleared out before full launch.
I think the rest was vultures combing through Discord logs and finding the devs don't much care for SJWs or somethin.
It caused enough of a hubbub that they were going to remove stuff, and donate to some LGBT cause, and then the other side threw a ruckus at censorship.
In the end, they're sticking to their guns, apparently, about 'Buy it or not. It's art.'